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gmatbull
In previous studies in which features of lichen communities were used as indicators of pollution, the
communities examined were located on tree stems 1–2 meters above ground, though lichen communities
on lower branches have shown to be more sensitive to changes in the air environment.

A. have shown to be
B. have since been shown to be
C. show themselves to be
D. are shown to be
E. are shown as

OA
after some discussions

IMO A.

D and E wrong because of active voice. we need passive.
"themselves in B makes no sense.
B is wrong because we have already used though in the sentence to show contrast so since is nt required
The last part reads ->
though lichen communities on lower branches have shown to be more sensitive to changes in the air environment, the
communities examined were located on tree stems 1–2 meters above ground
so A fits the bill.

what's the OA?
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The word previous studies indicate that there are going to be some later studies. While the communities happen to survive at 1 to 2 meters above ground ( which is the lower part of a tree), the later studies reveal that these communities do not survive at such low heights due to their sensitivity to changes in the air environment at those low levels. This is what the text means to say. The contrast reveled by the later studies are brought only in choice B, by using the time marker, sine.


Grammatically too, you do require a passive voice in the second part to be parallel to the passive voice in the first part. A and C are out for that reason. Among B, D and E, choice D and E miss the time-marker ‘since’ , thus leaving the question of when the contrast was established, wide open
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@Daagh,
supposing option B were "have BEEN shown to be", would it suffice?
In that case, B would be past perfect and passive too.

Meanwhile the correct response is Why must
"since"
be there?
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gmatbull
@Daagh,
supposing option B were "have BEEN shown to be", would it suffice?
In that case, B would be past perfect and passive too.

Meanwhile the correct response is Why must
"since"
be there?
'Since' is the most important part of the option B. As mentioned in earlier post, only this option fits the contrast presented in original sentence, and since helps to create that fit.
As per your question, whether u can omit since and then will it make sense.. ans should be no. If you read the entire sentence without since, it makes no logical sense.
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gmatbull
In previous studies in which features of lichen communities were used as indicators of pollution, the
communities examined were located on tree stems 1–2 meters above ground, though lichen communities
on lower branches have shown to be more sensitive to changes in the air environment.

A. have shown to be
B. have since been shown to be
C. show themselves to be
D. are shown to be
E. are shown as

OA
after some discussions


Oh!!! I eliminated C,D and E and was left with A and B because of the tense form used. Here we need a perfect tense as it is a previous study that is mentioned in the question.

Was not sure why should A be eliminated, although B sounds weird but the usage of 'since' is important to introduce the a contradiction and so the correct answer should be B
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In previous studies in which features of lichen communities were used as indicators of pollution, the communities examined were located on tree stems 1–2 meters above ground, though lichen communities on lower branches have shown to be more sensitive to changes in the air environment.

A. have shown to be - active voice - wrong
B. have since been shown to be - passive voice - correct
C. show themselves to be they con't show themselves
D. are shown to be - strange constraction since it is not clear shown by whom
E. are shown as - wrong meaning
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