The path of hurricanes can be carefully simulated, but computational errors, random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists can all combine to introduce more uncertainty than estimated.
(A) random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists can all combine to introduce more uncertainty than
(B) random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists can all combine to introduce more uncertainty than
they had
no reference of "they"(C) random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists can all combine to introduce more uncertainty than
they have
same as B(D) random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists
are the combined cause of the introduction of more uncertainty than
not parallel with "can"(E) random shifts in the flow of the jet stream, and the incomplete mathematical models used by meteorologists can combine to cause
the introduction of more uncertainty than
causes uncertainty not the introduction