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MA
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"A"...no other sentence convey the same meaning of "Why someting happened"
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"A" completes the sentence most naturally. This is an ongoing tradition...
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MA
Why painting the face and perfuming the body should have persisted throughout recorded history has been the subject of a substantial amount of anthropological and psychological speculation.

(A) Why painting the face and perfuming the body should have persisted
(B) How painting the face and perfuming the body could be persistent
(C) That painting the face and perfuming the body are persisting
(D) Painting the face and perfuming the body persisting
(E) The persisting of face painting and body perfuming


C, D create a sentence fragment. Persisting as a participle creates a sentence fragment.

E: Persisting modifies the wrong word.

B changes the meaning of the sentence. How Vs Why.

Therefore my AC is A. The "throughout" in A tells us that a present perfect tense is needed.
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MA
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OA is just A.



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