Although 35 Asian freshwater turtles
were initially placed in captivity in Miami’s Metro zoo in the summer of 1987, they had moved to the more temperate environs of the San Francisco Zoo in 1989 after zookeepers realised that the turtles could not survive the heat and humidity of southern Florida.
a. were initially placed in captivity in Miami’s Metro zoo in the summer of 1987, they had moved to the more temperate environs of the San Francisco Zoo in 1989
b. placed themselves in captivity initially in Miami’s Metro zoo in the summer of 1987, they were moved to 1989 in the more temperate environs of the San Francisco Zoo
c. were placed initially in Metro zoo of Miami in captivity in the summer of 1987, moving to the more temperate environs of the San Francisco Zoo in 1989
d. were placed initially in captivity in the Miami Metro zoo in the summer of 1987, they moved to the more temperate environs of the San Francisco Zoo in 1989
e. were initially placed in captivity in Miami’s Metro zoo in the summer of 1987, they were moved to the more temperate environs of the San Francisco Zoo in 1989