OE
The “but” that connects the two parts of the sentence indicates that the second part contrasts in some way with the frst part. In this case, the contrast is between the biologists’ agreement on a central fact (snakes’ descent from lizards) and a disagreement over the details (which have long been a matter of debate). Te words that best complete the blank, then, must somehow emphasize this disagreement. Of the choices, “competing” and “contending” best do so because they imply that the two theories are what the long-running debate has centered upon. Of the other choices, “conclusive” and “irrefutable” are roughly synonymous, but neither suggests a matter worthy of a long-running debate. “Dubious,” in addition to having no near-synonym among the choices, would also not explain why the emergence of the two theories triggered the debate. While “complex” might well describe a theory, it too has no obvious pair among the choices, and it also does not foreground biologists’ disagreement regarding the two theories.
Thus the correct answer is competing (Choice B) and contending (Choice E).