According to a wildlife reintroduction project, exactly four species will be reintroduced in one of three regions: Highlands, Wetlands, or Grasslands. Eight species are under consideration: Wolf, Bison, Otter, Lynx, Bear, Eagle, Elk, and Sheep. The reintroduction follows these rules:
• Eagle can be reintroduced only if the project is not in the Wetlands.
• Bear can be reintroduced only if Bison is also reintroduced.
• Bison can be reintroduced only if the project is in the Highlands or Wetlands.
• Wolf can be reintroduced only if the project is in the Highlands.
• Of the three species Otter, Lynx, and Sheep, exactly one may be reintroduced.
Based on the information above, and assuming Bear is reintroduced and the project is not in the Highlands, in the first column select the species that must also be reintroduced, and in the second column select the species that will not be reintroduced. Make only two selections, one in each column.