(B) The greatest road system built in the Americas prior to the arrival of Christopher
Columbus was the Incan highway, over 2,500 miles in length, and extended from northern Ecuador through Peru to southern Chile.
Structure:The greatest road system built in the Americas
(Subject)prior to the arrival of Christopher Columbus
(additional information telling us "when" the road was built)was the Incan highway
(Verb 1 - was ), over 2,500 miles in length,
(modifier modifying "highway")and extended from northern Ecuador through Peru to southern Chile.
(verb 2 - extended)Meaning:The sentence is talking about a road system that was built in America. This road system was once the greatest of its kind. It was called the Incant highway and it extended from northeast ecuador all the way to southern chile
(E) The greatest road system built in the Americas prior to the arrival of Christopher
Columbus, the Incan highway was over 2,500 miles long, extending from northern Ecuador through Peru to southern Chile.
Structure:The greatest road system built in the Americas prior to the arrival of Christopher Columbus,
(Modifier modifying "Incan highway")the Incan highway
(Subject)was over 2,500 miles long,
(Verb)extending from northern Ecuador through Peru to southern Chile.
(modifier modifying the previous clause - over 2500 miles long)
Meaning:The Incan highway was once the greatest road system built in America. It was over 2500 miles long. It extended from norther Ecuador to Chile