When testing the inactivated poliovirus vaccine that was licensed for a mass vaccination campaign in the United States in 1955, combined in it by the vaccine's creator, Jonas Salk, were three wild, virulent reference strains.(A) When testing the inactivated poliovirus vaccine that was licensed for a mass vaccination campaign in the United States in 1955, combined in it by the vaccine's creator, Jonas Salk, were three wild, virulent reference strains.
(B) Combining three wild, virulent reference strains, a test of the inactivated poliovirus vaccine that was licensed for a mass vaccination campaign in the United States in 1955 that Jonas Salk created.
(C) When the inactivated poliovirus vaccine created by Jonas Salk was licensed for a mass vaccination campaign in the United States in 1955, three wild, virulent reference strains had been combined in it.
(D) Three wild, virulent reference strains, combined in Jonas Salk's test of the inactivated poliovirus vaccine in 1955, that was licensed for a mass vaccination campaign.
(E) When Jonas Salk tested the inactivated poliovirus vaccine that was licensed for a mass vaccination campaign in the United States in 1955, he combined in it three wild, virulent reference strains.