I can see many questions over here but the answers available are limited so felt like sharing my view.
There were a lot of interactions between the two bases, which is how they stumbled across each other – although neither remembers the exact moment
In this question I've found in the non underlined part that "neither remembers the exact moment"
Which means there is clearly only one interaction then only this sentence make sense { Eliminate A,B}
A) There were a lot of interactions between the two bases, which is how they stumbled across each other – although neither remembers the exact moment-
Reason mentioned aboveB) There were a lot of interactions between the two bases, that is how they stumbled across each other – although neither remembers the exact moment
Reason Mentioned aboveC) There was a lot of interaction between the two bases, that is how they stumbled across each other – although neither remembers the exact moment
That should used in cases that are specific, So eliminate CD) There was a lot of interaction between the two bases, which is how they stumbled across each other – although neither remembers the exact moment
E) There was a lot of interaction among the two bases, which is how they stumbled across each other – although neither remember the exact moment { among is wrong usage for two bases}