Over the course of time, the mind had developed through the movements of the lips, tongue, jaw, and soft palate a sense of articulate speech, which has been integrated in the mind with rhythmical modulated sounds conveyed to the brain by the auditory nerves.
A) had developed through the movements of the lips, tongue, jaw, and soft palate a sense of articulate speech, which has -
no need of past perfect
B) has developed
a sense of articulate speech through the movements of the lips, tongue, jaw, and soft palate, which
have -
subject verb agreement errorC)
developed through the movements of the lips, tongue, jaw, and soft palate, and a sense of articulate speech has -
developed is participle - missing verbD) had developed a sense of articulate speech through the movements of the lips, tongue, jaw, and soft palate, which have -
no need of past perfectE) has developed through the movements of the lips, tongue, jaw, and soft palate a sense of articulate speech, which has -
correct, correct use of present perfect, and also corrects subject - verb agreement.
so E.