IMO B is the correct answer. Took me 2:20 minutes to decide between B and E. A very good question to brush up verb tense concepts.
Fifty years ago, ecologists studying wolf recolonization down from Canada into Washington State hypothesized that by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.
Let's cut the fluff first, we get- by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.
Meaning-
1- Wolves might reverse m overpopulation. How? by killing coyotes.
2- m overpop occured when- wolves were eliminated nearly 100 years back. (very important to understand this bit)
The glaring error in the original choice is the use of 'will' (future tense, also used when we definitely know that the action will occur in the future without fail). Ecologists have hypothesised. Therefore, even they can not be sure that the action they are suggesting will happen
definitely . Therefore, we must use 'would' instead of 'will'. 'Would' suggests that there is a probability (not 100%) that the action will occur.
Reject A, D.
B- No errors. Let's get back to this answer choice later.
C- "by killing coyotes, wolves would reverse the mesopredator overpopulation that had occurred when wolves had been"
E- "wolves would, by killing coyotes, reverse the mesopredator overpopulation that occurred when wolves had been"
When we see two actions that happened in the past, we get excited to use past perfect tense. And, in the excitement, we forget that there are two conditions that one must always keep in mind when using the perfect tense-
1- The two actions must not be independent of each other.
2- The use of past perfect is not 'compulsory' when some words or a phrase already provides the time context in the sentence. (for example, "nearly a century earlier" in the above sentence.)
Refer this-
https://gmatclub.com/blog/2016/05/past- ... orrection/A must read article.
That being said, let's see the errors in C, E.
C- 'that had occurred'- this phrase suggests that the action of occurring happened before some other
past action
not independent of occuring. There is no such
past action in the sentence. 'hypothesised' is an independent action, and has no link with 'had occurred'. Additionally, use of past perfect
continuous 'had been eliminated' is incorrect. Use of continuous tense means that the action of elimination is still happening. But this does not make sense, and is not the intended meaning. Choice E is incorrect for the same error.
That leaves us with B.