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Option C is the correct answer. The act of hypothesizing by the scientist the related action in given in simple past tense.
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Fifty years ago, ecologists studying wolf recolonization down from Canada into Washington State hypothesized that by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.

Timeline of events as per the stimulus. Wolves eliminated -> Mesopredator overpopulation occurs -> Ecologists (50 years back) hypothesise -> wolves will kill the coyotes and reverse the phenomenon

By creating a timeline of events in your head, it helps in determining the correct tense usage. So, let's see which answer choices use the best tenses and convey the intended meaning.

A) by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were
Ecologists hypothesised (past tense), correct tense to be used while expressing future from past- "would" not will

B) by killing coyotes, wolves would reverse the mesopredator overpopulation that occurred when wolves were
This answer choice is close by not perfect. By using simple past tense "when wolves were" it doesn't convey the correct timeline. It makes it seem that the coyote's overpopulation and the elimination of wolves were two simultaneous events. Eg: He cooked the meal, while she played the guitar. Both the events expressed in the simple past tense, occurring at the same time period in past. This is not the intended meaning. Correct usage will be of the past perfect tense.


C) by killing coyotes, wolves would reverse the mesopredator overpopulation that had occurred when wolves had been
This choice has both "would" and "had been" but it has an error in "had occurred", it distorts the timeline

D) wolves, by killing coyotes, will reverse the mesopredator overpopulation that had occurred when wolves were
This reverses the timeline. As per this choice. Timeline is Overpopulation occured -> wolves eliminated. This is not the correct timeline.

E) wolves would, by killing coyotes, reverse the mesopredator overpopulation that occurred when wolves had been
This answer choice presents the correct timeline by using the correct sequencing of events by using correct verb-tenses. Wolves would (by killing coyotes) reverse X that occurred when wolves had been eliminated a century ago. This precisely presents the sequence. Hence, the correct choice.
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This question is about verb tense:
A. will - occurring - were (eliminated: 50 years ago, cannot use will)
B. would - occurred - were (eliminated: 100 years earlier)
C. would - had occurred - had been (two latter events do not occur at the same time)
D. will - had occurred - were (eliminated: 50 years ago, cannot use will)
E. would - occurred - had been (correct answer)

Timeline: wolves eliminated from region -> mesopredator overpopulation -> wolves reverse
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IMO B is the correct answer. Took me 2:20 minutes to decide between B and E. A very good question to brush up verb tense concepts.

Fifty years ago, ecologists studying wolf recolonization down from Canada into Washington State hypothesized that by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.

Let's cut the fluff first, we get- by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.

Meaning-
1- Wolves might reverse m overpopulation. How? by killing coyotes.
2- m overpop occured when- wolves were eliminated nearly 100 years back. (very important to understand this bit)

The glaring error in the original choice is the use of 'will' (future tense, also used when we definitely know that the action will occur in the future without fail). Ecologists have hypothesised. Therefore, even they can not be sure that the action they are suggesting will happen definitely . Therefore, we must use 'would' instead of 'will'. 'Would' suggests that there is a probability (not 100%) that the action will occur.

Reject A, D.

B- No errors. Let's get back to this answer choice later.
C- "by killing coyotes, wolves would reverse the mesopredator overpopulation that had occurred when wolves had been"
E- "wolves would, by killing coyotes, reverse the mesopredator overpopulation that occurred when wolves had been"

When we see two actions that happened in the past, we get excited to use past perfect tense. And, in the excitement, we forget that there are two conditions that one must always keep in mind when using the perfect tense-
1- The two actions must not be independent of each other.
2- The use of past perfect is not 'compulsory' when some words or a phrase already provides the time context in the sentence. (for example, "nearly a century earlier" in the above sentence.)
Refer this- https://gmatclub.com/blog/2016/05/past- ... orrection/
A must read article.

That being said, let's see the errors in C, E.
C- 'that had occurred'- this phrase suggests that the action of occurring happened before some other past action not independent of occuring. There is no such past action in the sentence. 'hypothesised' is an independent action, and has no link with 'had occurred'. Additionally, use of past perfect continuous 'had been eliminated' is incorrect. Use of continuous tense means that the action of elimination is still happening. But this does not make sense, and is not the intended meaning. Choice E is incorrect for the same error.

That leaves us with B.
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Fifty years ago, ecologists studying wolf recolonization down from Canada into Washington State hypothesized that by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were eliminated from this region nearly a century earlier.

A) by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were -> we assumed something, so "will" is incorrect. we need "would".

B) by killing coyotes, wolves would reverse the mesopredator overpopulation that occurred when wolves were -> would is better to use. "When wolves were eliminated" gives a time frame as both action happened together. It is not the intended meaning. Incorrect.

C) by killing coyotes, wolves would reverse the mesopredator overpopulation that had occurred when wolves had been -> Are we saying, wolves are still eliminating from past. No! I think, it is an action completed, so the the mesopredator overpopulated. Incorrect.

D) wolves, by killing coyotes, will reverse the mesopredator overpopulation that had occurred when wolves were -> We can't say the mesopredator overpopulated first, then, wolves were eliminated. It is other way around. Incorrect.

E) wolves would, by killing coyotes, reverse the mesopredator overpopulation that occurred when wolves had been -> had been means an action completed in past, then another action "the mesopredator overpopulation" occurred. It makes sense. Let's keep it.

So, I think E. :)
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C is the correct answer as there are two activities happened in past in which the wolf elimination occurred earlier than overpopulation of coyotes

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Hi, In this sentence the timeline is clearly specified i.e the action of "hypothesized" happened 50 years earlier and the other 2 actions i.e "occurred & eliminated" took place a century earlier, so the use of past perfect tense is optional. So both B and C looks good to me. Can anyone please explain ?
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I think it supposed to be optio B. Can anyone explain me what is the problem with B?

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I would like help understanding why C is correct? I chose E. I was thrown off by "by killing coyotes"
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I picked C.

Here is my elimination method using splits of (1) will vs. would and (2) simple past tense vs. past perfect

A) by killing coyotes, wolves will reverse the mesopredator overpopulation occurring when wolves were future stated from point of the past should be expressed as a hypothetical as we're not sure it'll actually happen if we make a prediction from the past - it also says "hypothesized" so "would" is the right verb here

B) by killing coyotes, wolves would reverse the mesopredator overpopulation that occurred when wolves were there are two time points at play: (1) the time 50 years ago when ecologists hypothesized and in the question stem this is expressed as the past tense (2) the time when mesopredator population occurred because of the number of wolves going down which happened 100 years ago ("century") - In this sentence the elimination amd overpopulation happened at a time point earlier than scientists hypothesized so both "occur" and "is" should take the past perfect to denote this difference in timeline --> had occurred and had been

C) by killing coyotes, wolves would reverse the mesopredator overpopulation that had occurred when wolves had been right answer

D) wolves, by killing coyotes, will reverse the mesopredator overpopulation that had occurred when wolves were "would" should be used instead of "will"

E) wolves would, by killing coyotes, reverse the mesopredator overpopulation that occurred when wolves had beenoccurred is not in the past perfect as it should be to denote that it happened earlier than the scientists hypothesizing
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The official explanation is here.
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