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Subject - Verb

Presence of - remains, Eliminate A, B & E

Activity of studing began by archaeoligist, so who is appropriate after comma.
Eliminate D

My Answer C
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Quote:
The presence of mysterious paintings in a cave located outside Columbia, Missouri, remain puzzling to archaeologists, who in the mid-1980s began studying the images created by ancient residents of the Mississippi Valley.

Here, the subject is "Presence". So, we need "Remains". Presence is singular.
Eliminate A,B & E. (Subject Verb Agreement Error - Singular noun--> Singular verb, Plural noun--> Plural Verb)

We are down to C & D.
Comparing both, C is much more clear. The use of "Who" is referring to "archaeologists". This helps us to understand the sentence more clearly.
In the option D, beginning is modifying the preceding clause. ["Noun"+ "Comma + "Verb-ing"----> This modifies the clause, not the preceding noun]

Answer C
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In the sentence - The presence of mysterious paintings in a cave located outside Columbia, Missouri, remain puzzling to archaeologists.
the subject is "the presence" ,hence the correct verb will be "remains".

A) remain puzzling to archaeologists, who in the mid-1980s began studying
the subject is "the presence" ,hence the correct verb will be "remains".
B) remain puzzling to archaeologists, who in the mid-1980s began to study
the subject is "the presence" ,hence the correct verb will be "remains".
C) remains puzzling to archaeologists, who in the mid-1980s began studying
Correct S-V agreement. The archeologists began studying the painting.
D) remains puzzling to archaeologists, beginning in the mid-1980s to study
"beginning in the mid-1980s to study" acts as participle modifier and applies to the doer of the preceding clause. However,the presence of the painting did not begin the study.
E) remain puzzling to archaeologists, who in the mid-1980s would begin to study
the subject is "the presence" ,hence the correct verb will be "remains".

IMO C
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IMO C

The Subject is the presence , the plural noun of paintings is part of the prepositional phrase so dont get confused...
Hence Remain is wrong and we need Remains - A/B/E are out

Between C and D - D has ackward construct and C has both parts in past tence and meaning is intact hence C
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The presence of mysterious paintings in a cave located outside Columbia, Missouri, remain puzzling to archaeologists, who in the mid-1980s began studying the images created by ancient residents of the Mississippi Valley.

The subject is 'The presence' which is singular. Incorrect match of 'remain' (should be remains)
Cross out A,B,E
'who' correctly modifies the noun preceding who (archaeologists). 'The presence can't begin something' Cross out D.

A) remain puzzling to archaeologists, who in the mid-1980s began studying

B) remain puzzling to archaeologists, who in the mid-1980s began to study


C) remains puzzling to archaeologists, who in the mid-1980s began studying

D) remains puzzling to archaeologists, beginning in the mid-1980s to study

E) remain puzzling to archaeologists, who in the mid-1980s would begin to study
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