IanStewart
There's already a comma that shouldn't be there in the non-underlined portion, and I don't think there's any easy way, without bringing to the question outside information about US civil rights history, to decide between C and E. E is the correct answer here, but I don't much like it. It would be fine to write a sentence like "'Freedom Riders' traveled together on buses and forced police to protect protesters' rights". But when there's 30 words of intervening information between 'traveled' and 'forced', most readers are going to lose track of who is doing the "forced" part of the sentence. I think any writer who cares about clarity (which is the main concern of GMAT SC question writers) would restate the subject of the verb 'forced'. In short sentences, you can omit it. In long sentences, you really can't.
So I don't like E that much, but only E and C are free of grammar errors. And I can rule out C, because I know that the Supreme Court ruled that bus segregation was unconstitutional (that ruling wasn't enforced in some states, and the Freedom Riders' goal was to ensure that it was enforced). It was not true that there was an "unconstitutional Supreme Court ruling". So I know C is wrong because it contradicts historical fact. But I'm not supposed to use historical fact to choose between SC answers, and C is actually the best-written of the five answer choices, and it's certainly theoretically possible that the Supreme Court might make an unconstitutional ruling that protesters would challenge -- the meaning of C is not illogical. So I'm not really sure how a test taker is meant to answer this question without using historical information that test takers aren't expected to use.
I have no qualms with E. Long space between parallel items (including verbs) is very common on the test, and there is no other intervening conjugated verb that could be parallel with 'forced.' ('to test' is in the infinitive). The fact that some readers might lose track of something doesn't mean the structure is, objectively, unambiguous and clear.
(I took the real test two years ago. There was an SC question that was really long, and the underline was in the first part of the sentence. Every answer choice seemed fine, so I was confused, until I reminded myself I stopped reading halfway through the sentence... I checked the full sentence, and *miles* after the end of the underline, a conjunction built a parallel structure that made the answer obvious and unambiguous).
C, I agree that it requires too much civic/historical knowledge to understand that an 'unconstitutional supreme court ruling' doesn't make sense. I wonder if you could use the 'comma,--ing' modifier though?
"...who traveled together on buses into the southern states of Mississippi and Alabama to test the unconstitutional Supreme Court ruling on segregation of interstate buses, forcing federal police to protect..."
Hmm... Seems like 'forcing' would modify 'who traveled together on buses into southern states.'
Which... kind of makes sense to me? It does seem tough to get rid of C on any grounds except "I know the Supreme court ruling wasn't unconstitutional."