“while studying” suggests that the noun following the comma is the performer of the action “studying”.
A. a new kind of mutant genes was discovered by J. L. Scott, a discovery which led to greater understanding
1. “while studying… a new kind of mutant genes” – illogically suggest that mutant genes studied the genotype.
2. usually we need a comma before “which”.
B. . J. L. Scott discovered a new kind of mutant gene, a discovery led to more understanding
1. “a discovery led to more understanding” – illogically suggests that someone led the discovery to more understanding.
2. “more understand” implies that more people started to understand, while “greater understand” refers to the degree of understanding.
C. J. L. Scott discovered a new kind of mutant genes, and it led to more understanding
1. The only singular noun to which “it” can refer to is “a mew kind”. Implied meaning is illogical though: a new kind itself led to more understanding, not its discovery.
2. “more understand” implies that more people started to understand, while “greater understand” refers to the degree of understanding.
D. J. L. Scott, who discovered a new kind of mutant genes, led to greater understanding
1. “J. L. Scott led to greater understand” is not the intended meaning.
E. J. L. Scott discovered a new kind of mutant genes, a discovery that led to greater understanding
Bingo. The correct answer. All modifiers are used correctly.
Hence
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