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Salt deposits and moisture threaten to destroy the Mohenjo-Daro excavation in Pakistan, the
site of an ancient civilization that flourished at the same time as the civilizations …

The answer = “that” is required because if you omit it, the sentence will be a run-on sentence. But
consider another version of this sentence:

why is that required ? and is flourished modifier when used with that? how does it become run on sentence ? civilizations can't perform the action of being flourished themselves ?

Salt deposits and moisture threaten to destroy the Mohenjo-Daro excavation in Pakistan, the site of an
ancient civilization that was destroyed multiple times by flooding of Indus river…

Now in this sentence, “that” can be omitted (obviously along with “was”)

Here destroyed is being used as verb ?

Salt deposits and moisture threaten to destroy the Mohenjo-Daro excavation in Pakistan, the site of
an ancient civilization destroyed multiple times by flooding of Indus river…
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