In a cardboard representation of nucleotide subunits, Watson and Crick,
elaborating on other scientists’ theories which had failed to explain the nucleotide structures and pairings in their entirety,accurately modeled the double-helix DNA.
(A) elaborating on other scientists’ theories which had failed to explain the nucleotide structures and pairings in their entirety,
(B) elaborating on other scientists’ theories failed to explain the nucleotide structures and pairings in their entirety,
(C) elaborating on other scientists’ theories which have failed to explain the nucleotide structures and pairings in their entirety,
(D) elaborated on other scientists’ theories which fail to explain the nucleotide structures and pairings in their entirety,
(E) elaborated on other scientists’ theories which had failed to explain the nucleotide structures and pairings in their entirety,
Please explain which verb-ing form / verb-ed form will be used in the above answer?
Also explain the reasoning behind. Thanks in advance!
Please explain the rules for present participle in case a noun phrase is preceding?